There are probably other genes that influence the number and width of coloured bands on each hair and reduce or eliminate residual tabby markings on the chest, legs and sometimes tail. Yet other genes probably influence the sparkling appearance. In addition to the Agouti gene and Inhibitor (silver) gene, a Shaded cat may also have the Wideband gene (hypothetical) plus genes which influence the number and width of coloured bands on each hair and reduce or eliminate residual tabby markings on the chest, legs and sometimes tail. Yet other genes probably influence the sparkling appearance.
Some tipped and shaded silver shorthairs may have the ticked (Abyssinian-type) tabby gene in addition to the classic or mackerel tabby gene(s). This only shows up if matings of Shaded Silvers to Classic Tabbies unexpectedly produce ticked tabby kittens. Since ticked tabby masks out any other tabby pattern, it could only have been carried by the Shaded Silver parent. Tipped or Shaded Silver kittens born without a tabby pattern probably carry the ticked tabby gene, while those born with a discernible tabby pattern may lack the ticked tabby gene.
Breeder of Shaded Silver American Shorthairs, Carol W Johnson, suggested at least two modifier genes which affect ticked tabbies and which dissipate residual tabby markings in shaded shorthairs. She referred to them as Chaos and Confusion. A third gene. "Erase" was proposed by Cathy Galfo (working with Oriental Shorthairs) and appears to reduce or remove residual barring on the extremities. Currently, these names describe effects rather than actual genes.
Johnson noted that the each hair in a shaded cat differs in band number and width; ranging from solid colour and solid white through to multiply banded and singly banded. She termed this "Confusion" as it "unco-ordinated" the hair follicles to so that they produced different banding patterns to their neighbours. In contrast, Abyssinian-type ticked tabbies have even banding and relatively even colouration which stops abruptly at the belly (like a tide mark on a boat!). Shaded cats with high levels of Confusion (uneven band width) had a more mottled or "sparkling" appearance and a more gradual blending and fading of the colour from back to belly.
However, the confusion effect can be achieved with polygenes. Except for genes on X and Y chromosomes, each cell has 2 copies of each gene. The 2 copies might be identical or different and normally one is dominant to the other. Sometimes, e.g. when the genes are different but co-dominant, one copy is deactivated; this happens in foetal development. One cell might have activated the gene telling it to make 3 bands of colour while the cell next to it might have activated the slightly different copy of the same gene telling it to make 5 bands of colour. There might be a gene located elsewhere on the chromosome which, if switched on, override those genes entirely with an instruction to make a solid coloured hair! So while "Confusion" is a good name for the phenomenon on a visual level, a single Confusion gene seems unlikely.
In addition to Confusion, Johnson also hypothesised a rarer gene causing roan. In roan, solid white hairs are intermixed with normal hairs. This occurs in dogs (merle, roan) and horses (roan, flea-bitten grey). In Shaded shorthairs, "Roan" might result in Shaded Silvers so pale as be visually Chinchillas.
Johnson's "Chaos" gene further disrupts the striped pattern by abnormally mixing ticked hairs into normally solid coloured regions (and vice versa). This effect is visible in the modified tabby pattern of the Sokoke. An mechanism for this was described by Australian Mist breeder Truda Straede who suggested a gene which disrupted a normal tabby pattern into a "finely divided tabby pattern" (Striped and Spotted Cats). Straede had never seen the Sokoke, but predicted the patterns existence based on her work with "small pattern spotted tabbies" and "large pattern spotted tabbies". Chaos might eradicate residual necklaces and ghost striping in ticked tabbies and in Shaded cats.
Cathy Galfro proposed a separately inherited "Erase" gene, different to Confusion and Chaos, which removes residual markings from the neck, legs and tail of Shaded Oriental Shorthairs.